Charles Ashford, creator of the t-Virus, who enlisted the aid of the survivors in the city to recover his daughter. It was Cain who woke Alice and Matt Addison, now known as Nemesis, and let them loose in the city, hoping to test them against one another. This did not bother Cain in the slightest, as he believed that life was cheap and that in any case, no-one could logically have survived up to that point.
All efforts to contain the virus failed, and Umbrella sealed off the city in preparation for a nuclear strike. At Cain's direction, the Hive was unsealed, releasing the Undead and Lickers into the city, and resulting in a catastrophic outbreak of the t-Virus.
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